| Gram staining | 阴性 |
| Cell shape | 球状 |
| Cell length | 1.5-5 μm |
| Cell width | 0.2-0.3 μm |
| Spore formation | 不产芽孢 |
| Motility | None |
| Oxygen requirement | 微需氧 |
| Growth temperature | 37 °C |
| Optimal growth temperature | None |
| Growth pH range | None |
| Optimal growth pH | None |
| Growth salt concentration (w/v) | None |
| Optimal salt concentration (w/v) | None |
| Colony description | Colonies are round, 1–2 mm in diameter, raised, convex, smooth, and glistening. On moist media, colonies are flat, grayish, and spread in the direction of the streak. |
| Catalase reaction | None |
| Oxidase | None |
| V-P test | None |
| Methyl red test | None |